Nicolas Cage Grug Crood (voice) Emma Stone Eep Crood (voice) Ryan Reynolds Guy (voice) Peter Dinklage Phil Betterman (voice) Leslie Mann Hope Betterman (voice) Kelly Marie Tran Dawn Betterman (voice) Catherine Keener Ugga Crood (voice) Clark Duke Thunk Crood (voice) Randy Thom Sandy Crood (voice) Cloris Leachman Gran Crood (voice) Chris Sanders Belt (voice) Tara Strong Sash (voice) Melissa Disney Guy's Mother (voice) Artemis Pebdani Additional Voices (voice)